| Nantahala Capital Management, LLC | 2,820,838 | $30.2m | 0.5% |
| BlackRock, Inc. | 2,794,217 | $29.9m | 0.0% |
| AQR CAPITAL MANAGEMENT LLC | 967,859 | $10.4m | 0.0% |
| VANGUARD CAPITAL MANAGEMENT LLC | 833,596 | $8.9m | 0.0% |
| ORBIMED ADVISORS LLC | 759,155 | $8.1m | 0.2% |
| FEDERATED HERMES, INC. | 700,427 | $7.5m | 0.0% |
| VANGUARD PORTFOLIO MANAGEMENT LLC | 652,635 | $7.0m | 0.0% |
| MACKENZIE FINANCIAL CORP | 628,438 | $6.7m | 0.0% |
| Madison Avenue Partners, LP | 626,583 | $6.7m | 0.3% |
| Qube Research & Technologies Ltd | 608,837 | $6.5m | 0.0% |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 605,445 | $6.5m | 0.0% |
| STATE STREET CORP | 541,193 | $5.8m | 0.0% |
| GEODE CAPITAL MANAGEMENT, LLC | 510,841 | $5.5m | 0.0% |
| DIMENSIONAL FUND ADVISORS LP | 460,541 | $4.9m | 0.0% |
| CITADEL ADVISORS LLC | 396,098 | $4.2m | 0.0% |
| RENAISSANCE TECHNOLOGIES LLC | 359,086 | $3.8m | 0.0% |
| D. E. Shaw & Co., Inc. | 267,460 | $2.9m | 0.0% |
| BRIDGEWAY CAPITAL MANAGEMENT, LLC | 265,017 | $2.8m | 0.0% |
| Nuveen, LLC | 247,756 | $2.7m | 0.0% |
| ADAR1 Capital Management, LLC | 246,818 | $2.6m | 0.1% |
| TWO SIGMA INVESTMENTS, LP | 246,649 | $2.6m | 0.0% |
| ARROWSTREET CAPITAL, LIMITED PARTNERSHIP | 233,175 | $2.5m | 0.0% |
| MILLENNIUM MANAGEMENT LLC | 184,822 | $2.0m | 0.0% |
| GOLDMAN SACHS GROUP INC | 177,846 | $1.9m | 0.0% |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 158,712 | $1.7m | 0.0% |