| Portolan Capital Management, LLC | 1,299,256 | $48.7m | 1.8% |
| FMR LLC | 1,280,170 | $48.0m | 0.0% |
| BlackRock, Inc. | 1,183,443 | $44.3m | 0.0% |
| Allspring Global Investments Holdings, LLC | 1,072,916 | $37.8m | 0.1% |
| Driehaus Capital Management LLC | 846,354 | $31.7m | 0.2% |
| SUMMIT PARTNERS PUBLIC ASSET MANAGEMENT, LLC | 800,000 | $30.0m | 0.5% |
| DIMENSIONAL FUND ADVISORS LP | 782,276 | $29.3m | 0.0% |
| ROYCE & ASSOCIATES LP | 752,083 | $28.2m | 0.2% |
| VANGUARD CAPITAL MANAGEMENT LLC | 743,621 | $27.9m | 0.0% |
| First Eagle Investment Management, LLC | 739,150 | $27.7m | 0.0% |
| Boston Partners | 696,429 | $25.8m | 0.0% |
| Jupiter Topco LLC | 575,278 | $21.5m | 0.0% |
| BLAIR WILLIAM & CO/IL | 546,714 | $20.5m | 0.1% |
| WELLINGTON MANAGEMENT GROUP LLP | 477,583 | $17.9m | 0.0% |
| EVR Research LP | 441,908 | $16.6m | 8.2% |
| GEODE CAPITAL MANAGEMENT, LLC | 374,578 | $14.0m | 0.0% |
| JANE STREET GROUP, LLC | 356,151 | $13.3m | 0.0% |
| JPMORGAN CHASE & CO | 347,224 | $13.0m | 0.0% |
| KENNEDY CAPITAL MANAGEMENT LLC | 336,377 | $12.6m | 0.2% |
| Ophir Asset Management Pty Ltd | 326,699 | $12.2m | 1.0% |
| Granahan Investment Management, LLC | 324,108 | $12.1m | 0.4% |
| SILVERCREST ASSET MANAGEMENT GROUP LLC | 313,943 | $11.8m | 0.1% |
| STATE STREET CORP | 291,168 | $10.9m | 0.0% |
| Nuveen, LLC | 281,914 | $10.6m | 0.0% |
| BANK OF AMERICA CORP /DE/ | 276,317 | $10.4m | 0.0% |