| Granahan Investment Management, LLC | 12,393,660 | $186.4m | 6.6% |
| BlackRock, Inc. | 8,576,809 | $129.0m | 0.0% |
| D. E. Shaw & Co., Inc. | 5,289,385 | $79.6m | 0.0% |
| VANGUARD CAPITAL MANAGEMENT LLC | 4,508,442 | $67.8m | 0.0% |
| GILDER GAGNON HOWE & CO LLC | 4,381,990 | $65.9m | 0.7% |
| Park West Asset Management LLC | 3,536,810 | $53.2m | 3.6% |
| GOLDMAN SACHS GROUP INC | 3,222,042 | $48.5m | 0.0% |
| GEODE CAPITAL MANAGEMENT, LLC | 2,621,360 | $39.4m | 0.0% |
| STATE STREET CORP | 2,481,518 | $37.3m | 0.0% |
| Portolan Capital Management, LLC | 2,456,451 | $36.9m | 1.4% |
| VANGUARD PORTFOLIO MANAGEMENT LLC | 2,340,871 | $35.2m | 0.0% |
| Driehaus Capital Management LLC | 2,145,989 | $32.3m | 0.2% |
| Divisadero Street Capital Management, LP | 2,105,695 | $31.7m | 0.7% |
| DIMENSIONAL FUND ADVISORS LP | 1,665,626 | $25.0m | 0.0% |
| FRONTIER CAPITAL MANAGEMENT CO LLC | 1,547,453 | $23.3m | 0.2% |
| Jupiter Topco LLC | 1,461,364 | $22.0m | 0.0% |
| UBS Group AG | 1,423,495 | $21.4m | 0.0% |
| WELLINGTON MANAGEMENT GROUP LLP | 1,297,050 | $19.5m | 0.0% |
| FMR LLC | 1,184,434 | $17.8m | 0.0% |
| SILVERCREST ASSET MANAGEMENT GROUP LLC | 1,042,691 | $15.7m | 0.1% |
| ASHFORD CAPITAL MANAGEMENT INC | 976,320 | $14.7m | 1.6% |
| NORTHERN TRUST CORP | 867,794 | $13.1m | 0.0% |
| JPMORGAN CHASE & CO | 887,414 | $13.0m | 0.0% |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 849,852 | $12.8m | 0.0% |
| Pembroke Management, LTD | 823,718 | $12.4m | 0.9% |